Abstract
Introduction
As the global efforts to reduce carbon dioxide emissions associated with burning fossil fuels gain urgency, there is a growing sense that a range of industrial processes will need to be decarbonized in addition to power generation. The production of industrial gases provides a case in point. The predominant method of producing hydrogen, for instance, is through steam methane reforming which relies on natural gas as its energy feedstock and results in significant carbon dioxide emissions (Davis et al. 2018). A decarbonized alternative to steam methane reforming is to obtain hydrogen via a Power‐to‐Gas (PtG) process based on water electrolysis whereby electricity infused in water instantly splits the water molecule into oxygen and hydrogen. 1
The widely reported rapid decline in the cost of renewable electricity through wind or solar photovoltaic installations naturally raises the question as to whether a PtG system combined with an upstream renewable power source is becoming cost competitive with traditional, carbon‐intensive ways of producing hydrogen. A PtG system that is vertically integrated with a renewable power source will also have the financial advantage of avoiding the transaction cost typically reflected in the mark‐up of electricity prices for buying rather than selling electricity. 2
The intermittency of power generated from wind or solar photovoltaic sources presents a well‐known problem in balancing electricity supply and demand in real‐time. 3 One potential remedy suggested is to divert surplus energy from renewable power sources to the production of energy storing products like hydrogen. At the same time, though, the inherent intermittency of renewable power generation negatively affects the economics of a vertically integrated PtG system due to the high opportunity cost of “starving” the expensive PtG system at times of low output from the renewable power source; see Glenk and Reichelstein (2019). That bottleneck can be alleviated by allowing the PtG system to also draw power from the grid, though the resulting hydrogen will then become more expensive on account of facing the buying rather than the selling price of electricity. Furthermore, the hydrogen produced will be decarbonized only to the extent that grid power is decarbonized.
The front‐part of this study analyzes a generic model of a vertically integrated production system. The downstream unit requires an intermediate production input (e.g., power) that can be sourced from the external market, or alternatively from an upstream unit. The output of the upstream unit is inherently volatile as it fluctuates in an exogenous fashion across the hours of the year. Depending on the current market price of the intermediate input, the vertically integrated system can decide on a real‐time basis to what extent the intermediate input is sold on the external market or transferred to the downstream unit. In contrast to some of the recent work on production synergies, we capture volatility not by random shocks, but by predictable variations in both the level of the intermediate input produced and its market price (Hekimoglu et al. 2017, Kouvelis et al. 2018, de Véricourt and Gromb 2018).
The main question posed in the modeling part is whether a vertically integrated energy system exhibits
Our model analysis shows that the emergence of a synergistic investment value hinges critically on the two subsystems being sized optimally in terms of their relative capacity. 5 The need for this optimization reflects that in settings where the value of capacity investments is large relative to the annual operating costs, overall profitability is highly sensitive to trading off volatile revenue opportunities against idle capacity. We demonstrate that the optimal relative capacity size of two subsystems can be expressed in terms of a few aggregate variables. These comprise the life‐cycle cost of the intermediate and the final product, and the time‐averaged price and output levels, with the latter two averages adjusted by covariance terms that reflect the extent to which intertemporal variations in prices correlate with variations in output from the upstream unit.
The back‐end of our analysis calibrates our model in the context of PtG hydrogen production facilities that are co‐located with wind parks. We provide a numerical evaluation for vertically integrated energy systems in both Germany and Texas, two jurisdictions that have installed substantial amounts of wind power in recent years. On a stand‐alone basis, wind parks are currently unprofitable in Texas, though they entail positive NPVs in Germany, in large part due to public subsidies for renewable energy. The stand‐alone value of investments in PtG facilities depends on the attainable market price of hydrogen. For medium‐scale supply settings, hydrogen sales prices tend to be relatively high, making stand‐alone PtG facilities marginally profitable in both Germany and Texas. In contrast, such facilities entail negative NPVs in both jurisdictions relative to the lower prices associated with industrial‐scale hydrogen supply arrangements.
Since the integrated system generally experiences some operational gains from the avoided transaction costs that arise in the intermediate input market, one would expect a synergistic investment value to emerge if both wind power and hydrogen production are cost competitive (profitable) on their own. We confirm this for the setting of Germany and medium‐scale hydrogen supply. Conversely, it may intuitively appear difficult for the synergistic effect to be sufficiently large so as to outweigh stand‐alone losses if those occur in both subsystems. Nonetheless, we do identify such a synergistic investment value in the context of Texas where neither wind power nor hydrogen production is economically viable by itself.
An instructive metric for quantifying the gains from vertical integration is what we term the
We finally project likely improvements in the economics of combined energy systems that integrate wind power with hydrogen production. Several factors are likely to contribute to more robust synergistic investment values in the future. These include sustained price reductions for both wind turbines and PtG facilities as well as greater operational volatility in terms of fluctuating market prices for electricity. The latter trend is mainly a consequence of the trend towards time‐of‐use pricing. Overall, our projections indicate that even relative to the benchmark of the low hydrogen prices associated with large‐scale industrial supply, synergistic investment value for the integrated systems will widely emerge in both Texas and Germany within a decade. These projections take into account that the public support for wind energy, e.g., the production tax credit (PTC) available in the United States is scheduled to be phased out in the coming years.
For the specific application of wind power combined with hydrogen production, our numerical assessments point to more favorable economics than other recent studies (Ainscough et al. 2014, Bertuccioli et al. 2014, Felgenhauer and Hamacher 2015, Glenk and Reichelstein 2019). We attribute this difference to the fact that our calculations are based on subsystems that have been sized optimally, an aspect that is of first‐order importance when capacity investments account for a large share of overall production costs. In addition, our calculations take advantage of higher capacity utilization that results when both renewable and grid electricity are converted to hydrogen. Finally, our calculations reflect more recent cost and operational inputs for wind energy and PtG.
The remainder of the study is organized as follows. Section 2 develops the model framework for the identification of synergistic investment value in vertically integrated energy systems under conditions of operational volatility. Section 3 applies the model framework to PtG and wind energy. We first provide an assessment based on most recent data and then project likely changes in synergistic investment values for the coming decade. Section 5 concludes the paper. Supplemental materials such as proofs and data sources are provided in the Appendix.
Model Framework
Our model framework considers a vertically integrated energy system that comprises two interacting subsystems. For concreteness, we focus on a renewable energy source, like wind or solar power that is connected with a PtG facility. In our applications, the gas will be hydrogen that is produced via an electrolysis process. The setting in Figure 1 comprises four building blocks: the renewable energy source, the PtG facility, and external markets for both electricity and hydrogen.

Illustration of the Vertically Integrated Energy System [Color figure can be viewed at
We note that some ingredients of our model have been captured in earlier related work. For instance, in Hu et al. (2015), the PtG facility in our context is effectively replaced by an electricity customer who can obtain power from either the grid or his own renewable energy source. Alternatively, Kazaz (2004) studies an upstream unit that is an olive plantation facing fluctuating output levels and prices for olives, while the downstream unit is an olive press that converts olives into oil sold at a stable price. Similarly, it could be advantageous to combine an upstream pump for crude oil, that is subject to fluctuations in output and prices over time, with a downstream refinery system (Dong et al. 2014).
As a stand‐alone operating unit, the renewable energy source in Figure 1 can generate electricity that is sold on the open market at time‐varying prices. This stand‐alone subsystem is represented through the ellipse on the left. The boomerang‐shaped bubble on the right represents the stand‐alone PtG facility that can buy electricity from the open market to produce hydrogen sold at a time‐invariant price. Integration of the two subsystems enables the transfer of renewable power to the PtG facility (red arrow). Such integration will generally entail operational gains if the buying price of power exceeds the selling price faced by the renewable source. Any operational gains from vertical integration have to be traded‐off against the investment cost of capacity if one or both subsystems are not profitable on their own. Our analysis examines this trade‐off through the lens of an investor who seeks to maximize the NPV of the integrated energy system by optimally sizing the relative capacity of the two subsystems.
Contribution Margins
For given capacity investments, the integrated system shown in Figure 1 will seek to maximize the periodic contribution margin by optimizing the use of the available capacity in real time. The key variables in this optimization are the amount of power the renewable system produces at a particular point in time and the corresponding prices at which electricity can be bought and sold externally.
Let
Let
Given supply of electricity from either the external market or the internal renewable source, the
For a stand‐alone PtG system based entirely on electricity purchased on the open market, the contribution margin obtained at time
The contribution margin that can be attained from a vertically integrated system accumulates in four different phases that differ in terms of electricity prices and the conversion value of hydrogen. In Phase 1 of the diagram in Figure 2, both the buying and the selling electricity price exceed the conversion value:

Phase Diagram [Color figure can be viewed at
In Phase 2, the buying price exceeds the conversion value of hydrogen, which, in turn, exceeds the selling price:
In Phase 3, both electricity prices are non‐negative and less than the conversion value of gas:
Finally, in Phase 4, the buying price is negative and thus
The optimized contribution margin of a vertically integrated energy system at time
Lemma 1 shows that the contribution margin of a vertically integrated energy system can be expressed as the sum of the contribution margins of the two stand‐alone energy systems plus a third term that captures the economic interaction of the two subsystems. The term
Net Present Values
A vertically integrated energy system yields cash inflows in the form of optimized contribution margins. Such a system will create value if the discounted sum of the cash inflows collectively covers the initial cash outflow for capacity investments plus the subsequent periodic operating costs, including corporate income taxes. To identify conditions for the emergence of a synergistic investment value by the vertically integrated systems, it will prove useful to express the overall NPV in terms of unit costs and revenues. Specifically, we build on the definition of the
The LCOE aggregates all costs occurring over the lifetime of a power plant to deliver one unit of electricity output. The LCOE of a one kW facility can be expressed as:
Time‐of‐use electricity prices are frequently measured on an hourly basis. We denote by
To obtain the levelized capacity cost per kWh, the system price per kW is divided by the total discounted number of kWh that the system produces over its useful life:
Similar to the levelized cost of capacity, we define the levelized fixed operating cost per kWh as the total discounted fixed costs that are incurred over the lifetime of the facility divided by the levelization factor adjusted by the capacity factor.
It is readily verified that Δ is increasing and convex in the tax rate
Some countries, including the United States, grant subsidies in form of a tax credit for renewable energy production. For wind power, this takes the form of a PTC per kWh of electricity produced (U.S. Department of Energy 2016) as:
On the revenue side, the capacity factor,
In the terminology of Reichelstein and Sahoo (2015), the
The stand‐alone NPV of an intermittent power generation system is then given by:
For the hydrogen subsystem, our definition of the conversion value of hydrogen,
The NPV of a stand‐alone PtG facility can then be stated as:
Similar to the covariance between output and selling price for renewable electricity, we need to capture the co‐variation between hydrogen output and the price premium,
The NPV of the vertically integrated energy system of size
The first two terms of the NPV expression in Equation 24 reflect the value created by the two stand‐alone systems. The third term captures the time‐averaged synergistic gains. These gains are a function of both the average price premium (
An immediate consequence of Proposition 1 is that if both stand‐alone systems are profitable on their own, a vertically integrated energy system will generate synergies unless
Synergistic Investment Value
The vertically integrated system may exhibit synergistic investment value in each of the four possible scenarios that arise depending on whether the two stand‐alone systems are cost competitive on their own, or not. If indeed both subsystems are profitable on their own, one would expect a synergistic investment value that results from being able to by‐pass the electricity market and thereby avoiding the mark‐up between selling and buying prices for electricity. For completeness, we state the following formal result.
If both stand‐alone energy systems are cost competitive on their own, the vertically integrated energy system has synergistic investment value if and only if for some
Clearly, the inequality in Equation 26 can only hold during time intervals that correspond to Phases 2 or 3 in Figure 2. By Proposition 1, a synergistic investment value hinges entirely on
We next turn to the two mixed cases in terms of cost competitiveness of the two stand alone systems. Similar to the co‐variation factor
Suppose renewable power is cost competitive ( Suppose stand‐alone PtG is cost competitive, but renewable power generation is not. The vertically integrated energy system then has synergistic investment value if and only if:
Condition (28) in Proposition 2 states that the average price premium associated with PtG conversion must exceed the negative profit margin associated with the PtG system. The average revenue and life cycle cost associated with power generation are irrelevant since that activity will be undertaken regardless of the vertical integration decision. If the inequality in Equation 28 were to hold barely, the optimal PtG capacity would be small relative to the size of the renewable power source. To see this, suppose the capacity of the renewable power source is normalized to
An analogous argument verifies the necessity and sufficiency of the inequality in Equation 29 for the presence of a synergistic investment value if the PtG facility is profitable on its own, but renewable electricity generation is not. Holding the size of the electrolyzer fixed at
Overall, the

Linearity of the Optimal PtG Capacity Size [Color figure can be viewed at
In a hypothetical stationary environment where power prices and power generation are time‐invariant, we find if there is synergistic investment value in either one of the two scenarios identified in Proposition 2, the optimally sized vertically integrated system will be such that all renewable power is consumed internally. For scenario (i) in Proposition 2, (28) simplifies to
If neither stand‐alone subsystem is cost competitive on its own, an investor might still be willing to acquire a combination of the two subsystems provided the synergistic investment value more than compensates for the losses associated with the two stand‐alone systems. Figure 4 illustrates this possibility. Without loss of generality, we again anchor the size of the two subsystems, such that

Synergistic Investment Value if Neither Stand‐Alone Energy System is Cost Competitive on its Own [Color figure can be viewed at
If neither PtG nor intermittent renewable power is cost competitive on its own, a necessary and sufficient condition for a vertically integrated energy system to have synergistic investment value is that:
While the necessary and sufficient condition for synergies identified in Equation 34 is stated in terms of the endogenously optimized value
Suppose neither PtG nor intermittent renewable power is cost competitive on its own. The inequality:
The preceding claim is a direct consequence of Proposition 3 because, by construction, both
Application: Wind Energy and Power‐to‐Gas
Stand‐Alone Wind Energy
We now apply the preceding model framework to vertically integrated energy systems that combine wind power with PtG. Our numerical analysis focuses on Germany and Texas, two jurisdictions that have deployed considerable amounts of wind power in recent years. Wind energy naturally complements PtG as wind power tends to reach peak production levels at night when demand from the grid and electricity prices are relatively low (Reichelstein and Sahoo 2015, Wozabal et al. 2016). We base our initial calculations on 2017 data. Our data inputs are obtained from journal articles, industry reports, publicly available reports and interviews with industry sources (see the Appendix for a comprehensive list).
Wind energy is eligible for a federal PTC in the United States. It is paid per kWh of electricity generated (U.S. Department of Energy 2016). As shown in section 2, the PTC can be levelized and then effectively be subtracted from the LCOE. Beginning in 2017, Germany replaced its traditional fixed feed‐in premium for wind energy with a competitive auction system in which successful bidders are guaranteed a minimum price per kWh, with the government paying the difference between the successful bid and the actual revenue obtained from wind energy in the market place (EEG 2017). We refer to this difference as the Production Premium (PP). 12
Table 1 summarizes the calculation of the unit profit margin for wind energy in both jurisdictions. 13 The LCOE of wind energy amounts to 4.83 € ¢/kWh in Germany. The substantially lower LCOE 2.41 $ ¢/kWh in Texas reflects the impact of the PTC and, to a smaller extent, a higher capacity factor. The average selling prices of electricity amount to 3.46 € ¢/kWh and 2.44 $ ¢/kWh, respectively. The corresponding co‐variation coefficients of 0.87 and 0.93 indicate that prices tend to be below their average values during periods of above average wind output. We interpret the procurement auctions in Germany as competitive and therefore the profit margins are zero by construction. Thus, we infer the PP as the difference between the winning bids and the observed selling prices adjusted by the co‐variation coefficients. The estimates we obtain are corroborated by the observation that the range of observed winning bids (guaranteed selling prices) in 2017 was between 3.82 and 5.71 € ¢/kWh and our independent LCOE estimate is just about in the middle of that range.
Profit Margins for Wind Energy
Stand‐alone Power‐to‐Gas
As a producer of industry gases, a PtG facility in Germany is eligible to purchase electricity at the wholesale market price plus a relatively small markup for taxes, fees and levies. For Texas, we use the industrial rate offered by Austin Energy. Because of its grid connection, the PtG facility can also provide frequency control to the grid by rapidly absorbing excess electricity to balance supply and demand. Incorporating the revenues from frequency control into the price at which the facility can purchase electricity, the buying price of electricity averages 3.93 € ¢/kWh in Germany and 5.39 $ ¢/kWh in Texas (see the Appendix for details).
A PtG facility could be installed onsite or adjacent to a hydrogen customer. 14 The observed market prices for hydrogen are clustered in three segments that vary primarily with scale (volume) and purity. In Germany, prices for large‐scale supply amount on average to 2.0 €/kg, for medium‐scale to about 3.5 €/kg, and for small‐scale to at least 4.0 €/kg. In Texas, large‐scale hydrogen supply is priced at about 2.5 $/kg, while medium‐ and small‐scale are priced at about 4.0 $/kg or above 4.5 $/kg, respectively (Glenk and Reichelstein 2019).
Table 2 summarizes the calculation of the unit profit margin for PtG in both jurisdictions. The LFCH of PtG amounts to 2.36 € ¢/kWh in Germany and 2.22 $ ¢/kWh in Texas. For medium‐scale supply, the conversion premium of hydrogen amounts to 2.93 € ¢/kWh in Germany and 2.67 $ ¢/kWh in Texas, with corresponding profit margins of 0.57 € ¢/kWh and 0.44 $ ¢/kWh, respectively. For large‐scale hydrogen supply, the conversion premium equals 1.12 € ¢/kWh in Germany and 0.54 $ ¢/kWh in Texas and the corresponding profit margins are −1.24 € ¢/kWh and −1.69 $ ¢/kWh respectively. In terms of our model, we thus have the scenarios of the Corollary to Proposition 1 or Proposition 2 in Germany depending on the scale of hydrogen sales, while the setting in Texas corresponds to either Proposition 2 or Proposition 3.
Profit Margins for Power‐to‐Gas
Vertical Integration of Wind Energy and Power‐to‐Gas
The hydrogen prices shown in Table 2 for Texas and Germany show that our analysis spans the four possible scenarios that can arise in terms of the stand‐alone profitability of the two subsystems. Figure 5 indicates the presence or absence of a synergistic investment value for the vertically integrated PtG system. As one might expect, there is a synergistic investment value in Germany relative to the scenario of high hydrogen prices in the medium‐scale supply segment. Since both subsystems are profitable on their own in that scenario, the threshold for the presence of a synergistic investment value, that is, a conversion premium that is positive rather than zero (Corollary to Proposition 1), is indeed met.

Synergistic Investment Value of Vertically Integrated Wind Energy and PtG System [Color figure can be viewed at
Due to the relatively low hydrogen prices (large‐scale supply) in Germany, PtG exhibits a highly negative profit margin of −1.24 € ¢/kWh on its own. The synergistic price premium,
To quantify the synergistic investment value of an integrated wind energy and PtG system, it will be instructive to calculate the

Break‐Even Prices for Hydrogen Production [Color figure can be viewed at
Break‐even analysis can also quantify the value of giving the vertically integrated energy system access to buying electricity from the open market. Cutting off that supply branch would effectively yield a measure for the cost of renewable hydrogen, that is, hydrogen produced exclusively from wind energy. 16 Figure 6 shows the break‐even prices for renewable hydrogen as “renewable” prices. By construction, these prices must be higher than those of the vertically integrated system. The price difference is relatively large for Germany, indicating that in the current market environment access to the open electricity market is particularly important for the economics of hydrogen production.
To conclude this section, we solve for the optimal (relative) size of the PtG capacity for a given wind power facility the size of which has been normalized to 1 kW. The blue lines in Figure 7 display the NPV of the vertically integrated system as a function of the size of the PtG facility for alternative hydrogen prices ranging from 1.0 to 4.0 € or $ per kg. Red circles mark the optimal PtG capacity size for a particular hydrogen price. Circles at 0.0 kW indicate that no PtG capacity should be installed, while a red circle at 1.0 kW indicates that PtG is cost competitive on its own. As demonstrated in section 3, the NPV is always a single‐peaked function of

Optimal Power‐to‐Gas Capacity Size [Color figure can be viewed at
In comparison to other recent studies on the economics of hydrogen, our results point to generally lower hydrogen prices (Ainscough et al. 2014, Bertuccioli et al. 2014, Felgenhauer and Hamacher 2015). We attribute this discrepancy to several factors. Most importantly, our calculations are based on vertically integrated energy systems that are sized optimally for highly capital‐intensive capacity investments. In addition, our vertically integrated PtG facility is assumed to be connected to the grid and therefore obtains higher capacity utilization by converting renewable and grid electricity than it could achieve if it was to convert only renewable energy (Glenk and Reichelstein 2019). Finally, our calculations are based on most recent data reflecting the rapidly falling cost of producing wind energy as well as recent changes in the acquisition cost of electrolyzers.
Prospects for Synergistic Investment Value
The preceding numerical findings assess the economics of wind energy combined with PtG on the basis of recent data. Going forward, multiple trends appear to be underway that suggest further improvements in the economics of such vertically integrated energy systems. In this subsection, we incorporate the magnitude of these trends to track changes in the break‐even prices for hydrogen in future years. The break‐even hydrogen prices for a vertically integrated system reported in Figure 6 are the starting points of this trajectory.
Regarding the cost structure of wind energy, we follow Wiser et al. (2016) who project that the system prices for wind turbines will decline at a rate of 4.0% per year. At the same time, these authors project an increase in the average capacity factor at an annual rate of 0.7% per year. For the acquisition cost of electrolyzers, we rely on the regression results of Glenk and Reichelstein (2019), yielding an annual 4.77% decrease in the system price of PEM electrolyzers.
Our projections also assume that wind power in Germany and Texas will have a “driving role” in future changes of the selling prices of electricity in the wholesale market (Ketterer 2014, Paraschiv et al. 2014, Woo et al. 2011). Specifically, the difference between the LCOE in year
Figure 8 shows the trajectory of break‐even prices for hydrogen from a vertically integrated wind power and PtG system through 2030. Specifically, the hydrogen produced in this manner is projected to become cost competitive with industrial‐scale hydrogen supply, that is currently produced from fossil fuels, in the coming decade. The values shown by the solid line in Figure 8 assume an adjustment rate of

Trajectory of Future Hydrogen Break‐Even Prices [Color figure can be viewed at
Finally, we seek to capture the idea that further increases in renewable energy are likely to increase the variance in daily and seasonal electricity prices. As noted in section 2, higher operational volatility will generally tend to accentuate the synergistic investment value of a vertically integrated system. We incorporate the possibility of increased volatility in the selling price of electricity by assuming that
Conclusion
This study has examined the synergistic investment value of vertically integrated production systems. Synergies arise because of market imperfections for an intermediate input (electricity in our context) and because of operational volatility in the form of temporal fluctuations in output and prices. While vertically integrated systems will generally experience some synergistic benefit, we attribute a synergistic investment value only if a negative NPV for one or both of the stand‐alone systems is more than outweighed by the synergistic effect. In the context of an energy system that combines renewable energy with hydrogen production, we derive necessary and sufficient conditions for the presence of the synergistic investment value. These conditions can be stated in terms of life‐cycle unit costs and average prices adjusted for covariance terms that capture the extent to which price premia and output fluctuations are aligned across the hours of a typical year.
We rely on recent production price and cost data to assess the magnitude of synergistic effects in both Texas and Germany. Our empirical focus is on PtG facilities that can draw electricity either from the grid or internally from wind turbines. The policy support for renewable energy in Germany ensures that wind power is cost competitive on its own. We find that the emergence of a synergistic investment value in Germany hinges on the market price of hydrogen being above some break‐even value which is currently below the price paid for medium‐scale transactions, but above that obtained for industrial‐scale transactions.
Owing to the low wholesale prices of electricity in Texas, we find that, wind energy on its own is currently not cost competitive despite the PTC available to renewable energy in the United States. Nevertheless we find that the synergies between the two subsystems are sufficiently strong in Texas so that a vertically integrated energy system can create value, despite the fact that PtG facilities will also not be viable on their own.
While our numerical analysis is based on the most recent available data, several factors suggest a trend towards a more favorable economics for vertically integrated systems in the future. We base our forecast based on the combination of projected reductions in system prices for both wind turbines and electrolyzers as well as a general trend towards more volatility in electricity prices.
Our paper suggests several promising avenues for future research. With regard to the modeling part, it would be instructive to add stochastic shocks to prices and output. Such shocks are likely to increase the call option value of capacity investments, but it remains an open question whether additional volatility in the form of random shocks will lead to synergistic investment values for a broader range of circumstances. We also note that our framework has viewed hydrogen as a final product. An alternative and promising avenue is to view hydrogen also as a form of electricity storage. Provided the electrolyzer can also run in the “reverse direction,” hydrogen production coupled with reconversion to electricity may effectively compete with battery storage for electricity supply systems characterized by intermittent generation patterns.
