Abstract
How to Cite this Article
Usui, K., Ishikawa, M., Taya, S., & Kitaoka, A. (2026). Rotating dual-layered checkerboard illusion.
In everyday visual environments, physical continuity of objects is frequently obscured by occlusion or low contrast with their surroundings, resulting in physically continuous contours appearing as fragmented segments in retinal images. However, through motion of the observer or the objects, the visual system can perceive unified contours and shapes not detectable when static. This perceptual completion across space and time is called spatiotemporal boundary formation (SBF; Shipley & Kellman, 1994). In this paper, we introduce a novel visual illusion related to SBF.
When a 2 × 2 two-colored checkerboard square is superimposed on an identical background and rotated, its perceived shape undergoes dynamic changes, transitioning sequentially from a square to a rounded square, then to a circle, and back again (Movie 1 left). The perceived size also varies with shape, appearing largest with the square, smaller with the rounded square, and smallest with the circle (Figure 1). We refer to this phenomenon as the

Static version of RDCI.
Several visual illusions show changes in perceived shape and size linked to SBF in rotational motion. Among them, those creating a “breathing” impression are classified as “breathing illusions” (BI; Bruno, 2001). Under this definition, RDCI can be considered a new BI variant, though its stimulus configuration has distinctive features: in conventional BI displays, changes in visible edge length are determined by the aperture size (the slit between the occluders; see Movie 1 right), whereas in RDCI they are determined by the zero-contrast border between foreground and background. This gives RDCI a far larger visible-edge variation range than BI: in BI, it runs only from the aperture size

(A) RDCI static frames in several rotation angles (SR in parentheses). (B) Participants’ ratings vs. SR/rotation angle, and (C) ordinal probit model fit. Solid line and ribbon indicate the mean/fit with 95% CI. Dots indicate individual data points.
We considered whether visible edge proportion might determine the apparent shape of the RDCI. The ratio of visible to total (visible + illusory) edge length, called the
Two main accounts have been proposed for the BI. The first attributes it to
Interestingly, when the square element of the RDCI is translated, the size/shape changes similar to those seen during rotation can still be observed (Movie 2 left). This also might support the explanation based on the contour completion in BI, as it predicts that the illusion occurs regardless of motion type. However, it has been suggested that the BI weakens or disappears without rotational motion. For example, a translating square behind occluders typically appears as a nonrigid blob (Movie 2 right, “eccentric square” in Gerbino, 2017) or could appear as a rigid square only under certain conditions (Bruno, 2001, p. 545), neither showing the characteristic “breathing” impression.
Nevertheless, phenomenological differences in these translational versions may also support contour completion as the main mechanism of BI-family phenomena. In translational BI, visible edges appear asymmetrically (Figure 3B), and completion laws predict irregular contours (e.g., Gerbino, 2024). Although such asymmetric visible edges are also shown in translational RDCI (Figure 3A), the cross-shaped border formed by the white and black regions on the moving square remains constant in both size and shape, which may constrain the degrees of freedom of contour completion; for example, this may keep completed contours symmetric, and the perceived size may not fall below the size of the cross. In sum, while stimulus structure shapes how completion occurs, both phenomena can be understood as arising from the contour completion process.

Static frames of translational RDCI (A) and BI (B).
Although the present discussion has treated BI and RDCI as products of a common mechanism, their stimulus differences complicate direct comparison, leaving their precise relationship open for future study.
